Two critically endangered black rhinos have been welcomed to the Yorkshire Wildlife Park.
Jasper and Makibo were transferred from German zoos as part of a global breeding programme, which saw three-year-old rhinos Hodari and Dayo move from South Yorkshire to the Netherlands.
The heavyweight exchange involved months of planning at the wildlife park in Branton near Doncaster.
The incoming rhinos - of which there are fewer than 3,000 worldwide due to poaching - will make their first public appearances over Easter.
The exchange programme will see the Yorkshire Wildlife Park welcome a female black rhino named Najuma next year.