More than 100,000 London-bound rail commuters - a fifth of the overall total - have to stand at the busiest times of the morning rush-hour, according to Government figures.
Passengers on some morning and evening peak services are travelling on trains which are 60% over capacity on certain routes, the Department for Transport (DfT) figures showed.
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Inspectors found a culture of bullying and major problems with poor ambulance response times at the trust, which is the biggest in the UK.
More than a tenth of the Piccadilly Line train fleet is out of action because of emergency repairs to damaged wheels.
Rows of students stood and turned their backs before filing out of the lecture theatre.