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The UK's only female giant panda has been artificially inseminated after failed attempts to help her mate naturally.
Edinburgh Zoo bosses were hopeful that Tian Tian would mate with her intended partner Yang Guang this year.
But after her hormone levels started to fall quickly they turned to artificial insemination, using samples from the male.
Experts say they will not know for certain whether Tian Tian is pregnant until she gives birth, which could be in August or September.
The giant pandas at Edinburgh Zoo will be ready to mate within two weeks, keepers have revealed.
They hope Tian Tian and the zoo's male panda Yang Guang will produce a cub this year as the creatures both start to show the tell-tale signs that they are ready to breed.
There was disappointment last year when the pair did not mate, and although Tian Tian was artificially inseminated, she lost her foetus at late term.
Experts will watch Tian Tian's behaviour closely to gauge when her 36-hour breeding window begins.
Tian Tian (Sweetie) and Yang Guang (Sunshine) are the first giant pandas to live in the UK for 17 years. They arrived on loan from China in December 2011 and will remain at Edinburgh Zoo for a decade.
Zoo keepers have begun a daily monitoring of the UK's giant panda couple as mating season approaches.
Edinburgh Zoo hopes Tian Tian and Yang Guang will produce a cub this year as the creatures start to show the tell-tale signs they are ready to breed.
There was disappointment last year when the pair did not mate and though Tian Tian was artificially inseminated she lost her foetus at late term.
Panda reproduction is a notoriously tricky process, with females only ovulating once a year.
Edinburgh Zoo's giant panda Tian Tian is no longer pregnant, keepers have revealed.
Hopes were raised after Tian Tian showed signs of pregnancy, after she was artificially inseminated.
Tian Tian had been keeping keepers at Edinburgh Zoo guessing over her possible pregnancy since she was artificially inseminated in April.
In August, experts noted signs that she was pregnant and it was hoped a panda cub would be born by September and the panda enclosure was closed three weeks ago.
Chris West, Chief Executive Officer for the Royal Zoological Society of Scotland said:
“The panda enclosure will remain closed until the end of the week, in order to give Tian Tian time to get back into her routine and provide her keepers with the chance to recuperate after this long period of waiting.”
Experts at the Royal Zoological Society of Scotland believe giant panda Tian Tian "conceived and carried a foetus until late term, but then lost it".
– Chris West, Chief Executive Officer for the Royal Zoological Society of Scotland
We are all saddened by this turn ofevents after so many weeks of waiting.
Timings are difficult to pinpoint atthis moment, but we had a meeting this morning where Tian Tian’s behaviour and hormone results were reviewed and have come to the conclusion that it is very likely she has lost the pregnancy.
Giant panda Tian Tian is not expecting a cub, keepers at Edinburgh Zoo have confirmed.