The Danish zoo that caused international condemnation by putting down a healthy giraffe and dissecting it in public, has killed two lions and their two cubs to make way for a new male.
A spokesman told AFP: "Because of the lions pride, we had to euthanise those who were not old enough to fend for themselves."
The 10-month-old lions would have been killed by the new male lion "as soon as he got the chance", it said.
The Copenhagen Zoo went ahead with a plan to shoot and dismember a healthy giraffe in February and feed the 18-month-old animal's carcass to lions - an action the zoo said was in line with anti-inbreeding rules meant to ensure a healthy giraffe population.
A Danish animal activist says the killing of a healthy giraffe "should not have occurred" and was proof Copenhagen Zoo is "not the ethical institution that it wants to portray itself as being".
"Here we have a zoo which thinks that putting this giraffe down instead of thinking of alternatives is the best option," Stine Jensen, from Denmark's Organisation Against the Suffering of Animals, said.