South Yorkshire Police have confirmed that missing 14-year-old Lorna Vickerage has been found at Euston train station in London after being recognised by an off-duty police officer at 2.35pm.
John Bush, 35, has been arrested on suspicion of child abduction.
The force has thanked the media and public for their help in the case.
All air and sea ports have been put on alert for missing 14-year-old girl and 35-year-old John Bush.
Police say sightings suggest they are heading south from Yorkshire and that any move to leave the country would be "very foolish".
At a conference in Doncaster, South Yorkshire Police have said that Lorna Vickerage and John Bush reportedly tried to check into a hotel in Watford last night.
Police have received reports of sightings of a missing 14-year-old girl and 35-year-old John Bush in Leicestershire.
They were reported yesterday evening. Detectives believe the pair may have travelled to Melton Mowbray train station.
It's not yet known whether they boarded a train.
It is believed they were staying in accommodation in the Sproxton area.
The silver Ford Mondeo they were thought to be travelling in was recovered at around 6.40pm yesterday after being found in Sproxton. The car and accommodation are now being forensically examined.
Yesterday, police arrested a 35-year-old man from Askern, Doncaster on suspicion of child abduction and he remains in custody this morning.
Anyone with any information on their whereabouts is urged to call South Yorkshire Police on 101 quoting incident 609 of 10 June 2013.
Police searching for missing 14-year-old Lorna Vickerage have told ITV Daybreak that John Bush, who is wanted on suspicion of child abduction, is "known as a fraudster."
DCI Craig Robinson said: "There is no doubt he will use his skills as a fraudster to hide his identity and he will tell people [Lorna] is his daughter, when she's not."
Police looking for missing 14-year-old Lorna Vickerage have released photographs of a car they believe she might be travelling in with John Bush, a 35-year-old man wanted on suspicion of child abduction.
"Officers believe that they may be travelling in a silver Ford Mondeo, registration plate Y753 CSH, which John Bush had previously borrowed and not returned," a statement said.
DCI Craig Robinson, of South Yorkshire Police, said: “John Bush is wanted on suspicion of child abduction and officers have extensively searched the local area.
"We have alerted all forces and ports to assist with locating them, as we believe they may have now left the Doncaster area.
"Lorna has been known to go missing before, however not in these circumstances.”
A 35-year-old man from Askern has been arrested by police searching for missing 14-year-old Lorna Vickerage.
The man is not John Bush, the 35-year-old convicted fraudster, who is wanted on suspicion of child abduction.
The mother of Lorna Vickerage is "petrified" about her safety, police said. Detective Chief Inspector Craig Robinson of South Yorkshire Police said the 35-year-old man she is thought to be with has a history of fraud, and urged the 14-year-old to contact her family. Mr Robinson said:
Lorna, you're not in trouble. Your mum's petrified, she's worried to death, we're concerned for your safety. We don't know why you've gone with John and we don't know why John's taken you.
Let us know you're safe and well. Speak to John - tell him to hand himself in, and if you're listening, John we really do need to speak to you. The sooner you get this over with, the better.
Police in Doncaster have issued an appeal for a missing 14-year-old girl who is believed to be with a 35-year-old man.
Lorna Vickerage was last seen on Monday afternoon. South Yorkshire Police said:
"Lorna and John are thought to have been in the Highfields area over the last two days. Lorna was last seen at 2.30pm on Monday, 10 June at Outwood Academy in Adwick wearing her school uniform but it is unlikely that she will still be wearing this."