The number of post offices open on Sundays has reached 3,000 as the firm tries to keep up with changing demands, new figures show.
The milestone figure was reached when the Bergh Apton branch in Norfolk opened its doors - offering a garden centre, farm shop and restaurant as well as traditional postal services.
Our customers' needs are changing, and so are we. We intend to be the largest retail network in the UK open seven days a week.
Figures from The Post Office show the earliest branch opening time was 3.30am, while other branches close as late as midnight.
Three branches are open 24 hours a day.
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Staff in hundreds of Crown Post Offices will stage a fresh strike on November 20 in a long-running dispute over jobs, pay and branch closures, the Communication Workers Union said.
A Unite official has defended the union's decision for Post Office managers to go on strike.
Brian Scott said: "While we understand that the industrial action will cause inconvenience to the public, the ball is in the management's court - as it has been for the last 16 months - to reach a fair and equitable settlement.
"Unite is ready to have talks at anytime with the Post Office to resolve this matter, but the management has taken a very hardline stance against our members who have seen their living standards seriously eroded."
Hundreds of Crown Post Office managers will go on strike today in a row over pay.
Unite warned that the action will hit services at the larger branches, usually sited on high streets.
The union, which represents 900 managers, said its members have not received a pay rise since June 2011 and accused the Post Office of "dragging its feet."
Unite members will not undertake any overtime, will refuse to work any additional hours or do work not proper to their role in the organisation.
They will also refuse to travel outside of their working time, nor will they work in a location other than their normal place of employment.
This is an important day for Unite members in the Post Office.
They are keen to impress upon their employer their dissatisfaction with the failure to reach a pay agreement with the union and see this an opportunity to protest.
Post Office managers will stage their first industrial action in 34 years in a dispute over pay.
The Unite union, which represents 900 managers, said its members will ban overtime and work to rule for 24 hours.
The union accused the Post Office of "dragging its feet" for 16 months over a pay deal and warned of further industrial action if the dispute is not resolved.