Sam Burgess has been dropped from England's matchday 23 for Saturday's final World Cup group match against Uruguay at Manchester City Stadium.
Burgess has been omitted from the starting XV and bench as the hosts enter the death throes of a tournament that effectively ended for them with last Saturday's 33-13 defeat by Australia.
In a possible glimpse into England's future backline, George Ford has been recalled at fly-half, Owen Farrell moves to inside centre and rising star Henry Slade will win his second cap in the number 13 jersey.
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England and Wales will go head to head at Twickenham tonight in a sporting showdown that could influence which team will go through to the Rugby World Cup's knockout stages.
The potentially crucial match will see the long time rivals playing to secure their places at the top of the group table ahead of other Pool A opponents Australia, Fiji and Uruguay, as they play their second games of this year's contest.
Ahead of tonight's 8pm kick-of England coach Graham Rowntree said he believed Wales would be coming into the match with "doubts in their minds" after losing the last two games they have played against England.
But Wales fly-half Dan Biggar has insisted Wales will arrive at Twickenham confident and ready for battle as they "are fully aware of what is at stake."
Scottish and Japanese rugby fans have said they are both "nervous" and "super-excited" ahead of the World Cup clash between Scotland and Japan as one supporter called for a "miracle".
Wales have beaten underdogs Uruguay 54-9 in the Millennium Stadium, Cardiff.
Uruguay started well, scoring off three penalties, but Wales romped to victory with a total of eight tries.
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Japan has achieved a shock 34-32 win over South Africa in the Rugby World Cup.
Japan's victory in the Pool B match played in the Amex Stadium in Brighton came via a try in the 84th-minute.
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Thousands of fans from around the world gathered in a multi-coloured hoard to watch on big screens as England and Fiji kicked off the Rugby World Cup.
ITV News' Peter Smith reports:
As England took on Southern Hemisphere side Fiji at Twickenham, fans from both countries were joined by others from as far afield as Argentina, South Africa and Australia at a Fanzone in nearby Richmond Park as well as other zones around the country.
England have made a successful start to their Rugby World Cup campaign, beating Fiji, 35-11, at Twickenham.
However, England's less-than-convincing display will have left Australia and Wales scenting the hosts' blood.
Two Mike Brown tries, a penalty try and a last-gasp Billy Vunipola score saw England prevail, but Stuart Lancaster's men will need to improve dramatically ahead of next week's clash with Wales to maintain hopes of advancing from Pool A ahead of their neighbours and the Wallabies.
Fiji were a nuisance and, but for missed kicks, might have pushed England closer, but instead had to settle for Nemani Nadolo's try and two penalties.
As 86,000 fans enjoyed watching England take on Fiji at Twickenham tonight, thousands more have taken in the action at fan zones around the country.
And those in south London couldn't get much closer to the action when they packed into the Old Deer Park in Richmond just down the road from rugby HQ.
England and Fijian fans rubbed shoulders with others from as far afield as Argentina, South Africa and Australia at the Fanzone as they nervously watched the opening match in this year's tournament.
A sea of white, blue, yellow and red began filling the large site which provided pints of beer, hot food, fun fair rides, a practice rugby pitch and undercover big screens for the excited fans.
The Rugby World Cup will see 20 nations from around the world compete at venues across England and Wales, and around 2.3 million of the 2.45 million tickets on offer have already been sold.