Mike Tindall will play his 74th test for England on Saturday after being recalled for the Rugby World Cup clash with Romania.
Mark Cueto will also make his first appearance of the tournament as one of seven changes to the England team.
Cueto replaces Delon Armitage on the left wing, Jonny Wilkinson comes in for Toby Flood at fly-half and veteran Mike Tindall replaces Shontayne Hape at outside centre.
The pack has four changes with Steve Thompson in for Dylan Hartley at hooker, Alex Corbisiero in for the injured Sam Stevens at prop, Louis Deacon replacing Simon Shaw at lock, and Tom Croft replacing Tom Wood on the blindside flank.
Tindall fronted the media with coach Martin Johnson when the team to play Romania in Dunedin was announced, and the focus immediately switched to Tindall's big night out in Queenstown.
The 32-year-old Tindall was thrust into the spotlight after the release of video footage last week which showed him being kissed on the head by an unidentified woman ahead of England’s pool B win against Georgia.
The images sparked worldwide media attention as Tindall is married to Zara Phillips, the granddaughter of Queen Elizabeth II.
After the English media hounded Tindall about the controversy, Johnson took control and made it clear that any questions had to be related to the weekend's game.
"We've put it (the Tindall controversy) to bed," said Johnson. "We're getting on with playing so it's not an issue."
"We're looking forward to a big game this weekend. We had that conversation (about Tindall) last week, we've played a game since then and we've moved on."
Tindall plays for Gloucester and made his England debut in 2000.
He and Johnson, who was captain, were part of England's World Cup triumph in Australia in 2003.
Tindall played solidly against Argentina in the tournament opener but was rested for the match against Georgia last week.
His replacement, Shontayne Hape, scored two tries against Georgia but Tindall has been restored to the No.12 jersey against Romania.
"You ought to feel pressure all the time when you're playing for England," he said. "That's the nature of international rugby."
"The senior players among us have been around for a long time and you understand the frustration of the coaches when things you talked about all week at training don't happen on the playing field," he said.
"You can have a quiet word in people's ears about what's needed and hope that it helps.
"This week's training has been a big step up from where we've been and hopefully that will show on Saturday."
"There has been a bit of frustration. We talked about being relentless against Georgia but we weren't. We tried to skip some of the processes we needed to go through it we were going to score tries but it doesn't work like that."
"The frustration was that we made off-loads which weren't on, and conceded penalties and didn't do the things we'd practised all week."
All the England squad should have had game time after the weekend with front-rowers Lee Mears and David Wilson – the only players not to have seen action at the tournament – on the bench against Romania.
"We just need to go out and give ourselves a better chance," added Johnson. "We need to cut down on the penalties, cut down on the turnovers. If we do that, we feel we can be a lot better than we were last week.
"There has been a lot of honesty from the players since then. If we're satisfied with how we played against Georgia, we're not going to be in this tournament very long. This week's all about wanting to improve and to cut those errors out.
"Romania are strong in the set piece, strong in scrummaging and very strong ball carriers. It's our last game in Dunedin, it's a cracking place to play and we're all looking forward to getting back on the field. We've got to do everything we can to be as good as we can."
Romania, meanwhile, have made wholesale changes to their starting line-up with only four players - lock Cristian Petre, number eight Ovidiu Tonita, scum-half Lucian Sirbu and centre Ionel Cazan – retained from their loss to Argentina.
Inspirational captain Marius Tincu and several other leading players have been named on the bench as Romania juggle their resources to play England and Georgia in the space of four days.
"We trust this team," said coach Romeo Gontineac. "We trusted them before the World Cup so we trust them now as well.
"We want to share the playing time. A lot of players have played 80 minutes and we want everyone to enjoy the World Cup.
"We want these players to prove that they are good enough to play even against England, who are one of the best teams in the world.
"We want to achieve what we did in the first game, against Scotland, and to prepare for a very good game against Georgia next Wednesday."
England team: Ben Foden; Chris Ashton, Manu Tuilagi, Mike Tindall, Mark Cueto; Jonny Wilkinson, Ben Youngs; Alex Corbisiero, Steve Thompson, Dan Cole, Louis Deacon, Tom Palmer, Tom Croft, Lewis Moody (capt), James Haskell.
Replacements: Lee Mears, David Wilson, Simon Shaw, Tom Wood, Richard Wigglesworth, Toby Flood, Delon Armitage.
Romania team: Florin Adrian Vlaicu; Stefan Eugen Ciuntu, Ionel Cazan, Iulian Dumitras, Adrian Marian Apostol; Marin Danut Dumbrava, Lucian Mihai Sirbu; Nicolae Nere, Bogdan Zebega Suman, Silviu Florea, Valentin Poparlan, Cristian Constantin Petre
(captain), Sandu Stelian Burcea, Cosmin Aurel Ratiu, Ovidiu Tonita.
Replacements: Marius Tincu, Paulica Ion, Mihai Macovei, Daniel Gabriel Ianus, Valentin Nicolae Calafeteanu, Csaba Minya Gal, Catalin Marius Nicolae.