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Ed Miliband and Nicola Sturgeon are "at the altar, and preparing to sign the register" on a post-election deal, according to Scottish Conservative leader Ruth Davidson.
In a letter to party supporters she said: "In the last few days, we've now seen hard evidence that this nightmare scenario is now all too real.
"Another poll of Scottish seats on Friday showed that Scottish Labour is now a sinking ship, and is losing voters to the SNP. In truth, there is no way back for them.
"Their weakness means that one thing has become near inevitable: if Ed Miliband is in the position to form a government on May 8th, he will need the SNP's support to push him over the line.
"I said at the turn of the year that Ed Miliband and Nicola Sturgeon were halfway down the aisle towards a deal.
"The truth is that they are now at the altar, and preparing to sign the register. It's the only way Labour can get in to power."
When asked again today whether Labour would enter a formal coalition with the Scottish Nationalists to form a government, Ed Miliband repeated that it wouldn't happen.
However, the Labour leader did stop short of ruling out some sort of post-election pact, should his party fail to win a majority.
Meanwhile, SNP leader Nicola Sturgeon repeated her call for Labour to join forces, saying that only together could they lock the Tories out of Downing Street.
ITV News Deputy Political Editor Chris Ship reports:
Nick Clegg says he did not watch last night's TV debate, preferring to go to the pub in his Sheffield Hallam constituency instead.
Speaking during a visit to the Gordon seat in Aberdeenshire, the Liberal Democrat said:
Ed Miliband has accused David Cameron of trying to "duck, weave and dive" his way back into power, claiming he was "gutless" for refusing to take part in a head-to-head showdown.
"I'm offering him a debate about leadership. I'm offering him a one-to-one debate about the future of the country," the Labour leader said.
"I have been saying week after week, anytime, anywhere, any place."
Miliband joked that he knew the real reason behind Cameron's absence at lat night's debate, saing, "I hear he was washing his hair".
Ed Miliband insisted his message to Nicola Sturgeon is "thanks but no thanks" following her offer of a vote-by-vote deal to help him become Prime Minister.
"Vote Labour for a Labour government," Miliband told an audience of young people in Lincoln today.
"I think lots and lots of people are jumping way ahead here. I'm working for a majority Labour government," he said.
"I believe we can win a majority Labour government. Let's actually see what happens on May 7."
Nicola Sturgeon has said people across the UK will not forgive Ed Miliband if he turns down the chance to lock the Conservatives out of Downing Street.
Speaking on the campaign trail in Ayrshire today, Sturgeon said: "If there is an anti-Tory majority in the House of Commons after the election, then the only way David Cameron gets back into Downing Street is if Ed Miliband stands back and lets him back into Downing Street.
"If Ed Miliband and Labour do that, they will not be forgiven, certainly in Scotland and I suspect in large parts of the UK as well."
"It was very clear last night that on its own, Labour is not going to be bold enough to offer a real alternative to the Tories," she added.
David Cameron has claimed he was not invited to last night's televised leaders debate.
Asked by ITV News' James Mates why he had "ducked out" of the event, the Prime Minister said, "Last night was a 'challengers' debate'."
"It was the idea of the broadcasters to have a debate of the opposition parties, a debate to which Nick Clegg and I weren't invited," he continued.
Cameron was heavily criticised after he refused to sign up to the broadcasters' original plan of three debates.
David Cameron has warned there would be borrowing and taxes if Ed Miliband does a deal with the SNP to become Prime Minister.
The Tory leader said Miliband had failed to rule out a vote-by-vote deal with Nicola Sturgeon after she offered an olive branch during last night's "challengers' debate".
Cameron wrote on Twitter:
People watching last night's "challengers' debate" were most impressed by Ed Miliband and Nigel Farage, the ITV News/YouGov daily poll suggests.
Miliband and Farage topped the table with 15% each, with the SNP's Nicola Sturgeon a close third, according to a YouGov representative sample using the ITV News daily poll app.
Latest ITV News reports
The Ukip leader said he did not "lose my rag" when he attacked the audience during last night's televised debate for their "left-wing bias."
We watched undecided voters as they watched the debate hoping it might help them make up their minds.